Date/Place of Birth: 22nd October 1932
University education: 1950 - 1957
Natural Sciences, majoring in zoology, Free University of
Berlin; 1957 Dr. rer. nat.; title of dissertation:
"Verhaltensstudien am Trauerschnäpper" (Behaviour studies
of the Pied Flycatcher), Suppl. Z. Tierpsychol. No. 3,
Summer 1957 Research associate at the
Federal Bird Observatory of the Land Baden-Württemberg at
1957 - 1964 Research
associate at the Max-Planck-Institute for Behavioural
Physiology at Seewiesen (Department Lorenz).
1964 - 1967 Assistant
Professor at the Department of Zoology, Tübingen
Habilitation at Department of Biology,
Ruhr-Universität Bochum; ´Dozent´ and founder of the
Animal Behaviour Research Group (´Arbeitsgruppe für
Verhaltensforschung´), Faculty of Biology.
1950 - Comparative behaviour
research and population ecology of songbirds, with
emphasis on visual mechanisms (enemy recognition in birds,
prey recognition in reptiles), the functional organization
of complex activities, ethometry. Research trips to
Southern Macedonia (1958), Spain (1960), Galapagos
(1962/63), Jamaica (1969), Panama (1979), Tonga (1990),
Fiji (1990/91), and Philippines (1993
1963 - studies on the
behavioural and morphological phenomorphism of
neo-tropical hawkmoths and the measurement of the
selective coefficient of larval phenomorphs; stochastic
determination of intra-specific behavioural variation of a
Jamaican lizard. Sociobiology and life history of parental
care in songbirds.
- Doctoral Dissertation (1959, Parey, Berlin, Hamburg)
- The Ethology of Predation (1976, Springer). A review of
the ethology and behavioural ecology of predator hunting
Physiological practical; excursion in marine biology to
Italy; courses, seminars and practicals in ´Behaviour´;
further courses in ´Zoology for Psychologists´;
´Introduction to quantitative evolutionary research ´;
´Behavioural ecology: empirics and modelling´.
Public education, lectures and seminars, e.g. ´Models in
biology´ together with K. STEINBUCH on the European Forum
at Alpbach (Austria).
Supervision of 12 dissertations and ca. 40 High School
teacher and master's theses.
1968 - 1972 Representative of the Deans of the
mathematical and natural sciences of the Ruhr-University
at the West German Mathematical and Natural Sciences
Faculties Meeting. Member of the coordination committee
during the initial build-up of the Ruhr-University as well
as member of University Parliament.
1968 – 1997 Faculty member of the Department of Biology,
Ruhr-University; various activities in councils and
1965 - 1977 serving on the Scientific Advisory
Committee of the German Ornithologists´ Society (´DO-G´),
1973 President of that Committee
1974 President of the Society´s Research
1966 - 1972 Scientific Advisory Committee of
the Charles Darwin Foundation for the Galapagos Isles
1967 - Member of WWF (Worldwide Fund for
1970 Elected member of the American Society of
1970 Member of German Conservation
Organization (´Bund für Umwelt- und Naturschutz
1973 - 1985 Editor of ´Zeitschrift für
Tierpsychologie´, now ´Ethology´.
1980 - 1996 Co-editor of ´Bird Behaviour´.
1980 - 1998 Election member of the
International Ethological Committee.
1987 - 1990 President of the Ethological
Society (´Ethologische Gesellschaft´).
1986 - International Society for Behavioural
1988 - 1998 European Society for Evolutionary Biology.
Plenary session lectures on meetings 1970 - 1991 of the
following societal or other institutions: Association for
the Study of Animal Behaviour (twice); German
Ornithological Society; German Zoological Society, Sociéta
Italiana di Etologia, Association of German Biologists,
European Society for Evolutionary Biology, International
Ethological Conferences (each one, the latter twice).
1991 - Member of the Society for Conservation
1992 Tinbergen lecture of the
Association for the Study of Animal Behaviour.
1994 Member International Ornithological
1995 Ornithologists´ Prize 1995 of the
German Ornithologists´ Society.
1995 - Founder and Director of Philippine
Endemic Species Conservation Project
1996 Research Award
towards studies in tropical biology of the Smithsonian
Institution in Central America.
1998 Election Member of IUCN Survival Service
Commission and its Re-introduction Specialist Group.
1999 - Election Editorial Board ‘Silliman Journal’
2000 Honorary member of Ethological
Society (Ethologische Gesellschaft)
2000 - 2001 President and co-founder of
Bioresource Conservation Trust for the Philippines
(BioCon), Pandan, Antique, Philippines
2001 - 2003 Board Member of BioCon
2001 - Chair for Biodiversity, first in the ASEAN
states, assigned by EU ASEAN Regional Centre for
2011 Renewal of the Philippine conservation
projerct and it´s umbrella NGO under new names with
slightly updated aims